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Smithsonian National Museum of Natural History
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Department of Invertebrate Zoology

Nemertean, Tubulanus polymorphus

Tubulanus rhabdotus (Palaeonemertea)

bar Norenburg image

Jon L. NorenburgResearch Zoologist

  • Phone: 202-633-1783
  • Fax: 202-633-0182
  • E-mail:

  • Mailing Address:
    Smithsonian Institution
    PO Box 37012, MRC 163
    Washington, DC 20013-7012

  • Courier Delivery Address:
    Smithsonian Institution
    National Museum of Natural History
    10th and Constitution Ave, NW
    Washington, DC 20560-0163

    Responsibilities: Research Zoologist and Curator of Nemertea

    Education: B.A. (1973), M.S. (1976) Acadia University; Ph.D. (1983) Northeastern University.

    Research Interests: Nemertean worms, worldwide – morphological and molecular phylogeny, taxonomy, phylogeography, biogeography and functional anatomy; soft‐bodied marine interstitial fauna, worldwide – biology, taxonomy and zoogeography.

    Professional Activities:
    Full Curator, Nemertea for Encyclopedia of Life
    Editor, Nemertea for World Register of Marine Species

    Recent Publications:

    Mendes, Cecili B., Norenburg, Jon L., Solferini, Vera N. and Andrade, Sónia C. S. 2018. Hidden diversity: Phylogeography of genus Ototyphlonemertes Diesing, 1863 (Ototyphlonemertidae: Hoplonemertea) reveals cryptic species and high diversity in Chilean populations. PloS One, 13(4): 1-22. doi:10.1371/journal.pone.0195833

    Scarpa, Fabio, Cossu, Piero, Delogu, Valentina, Lai, Tiziana, Sanna, Daria, Leasi, Francesca, Norenburg, Jon L., Curini-Galletti, Marco and Casu, Marco 2017. Molecular support for morphology-based family-rank taxa: The contrasting cases of two families of Proseriata (Platyhelminthes). Zoologica Scripta, 46(6): 753-766. doi:10.1111/zsc.12251

    Leasi, Francesca, da Silva Andrade, Sonia C. and Norenburg, Jon L. 2016. At least some meiofaunal species are not everywhere. Indication of geographic, ecological and geological barriers affecting the dispersion of species of Ototyphlonemertes (Nemertea, Hoplonemertea). Molecular Ecology, 25: 1381-1397. doi:10.1111/mec.13568

    Ament-Velásquez, S. L., Figuet, E., Ballenghien, M., Zattara, Eduardo E., Norenburg, Jon L., Fernández-Álvarez, F. A., Bierne, J., Bierne, N. and Galtier, N. 2016. Population genomics of sexual and asexual lineages in fissiparous ribbon worms (Lineus, Nemertea): hybridization, polyploidy and the Meselson effect. Molecular ecology, 25(14): 3356-3369. doi:10.1111/mec.13717

    Mendes, Cecili B., Matthews-Cascon, Helena and Norenburg, Jon L. 2016. New records of ribbon worms (Nemertea) from Ceará, Northeast Brazil. Zootaxa, 4061(2): 146-156. doi:10.11646/zootaxa.4061.2.4

    Leasi, Francesca, Gaynus, Camille, Mahardini, Angka, Moore, Tiara N., Norenburg, Jon L. and Barber, Paul H. 2016. Spatial and ecologic distribution of neglected microinvertebrate communities across endangered ecosystems: meiofauna in Bali (Indonesia). Marine Ecology-an Evolutionary Perspective, 37(5): 970-987. doi:10.1111/maec.12305

    Scarpa, Fabio, Cossu, Piero, Sanna, Daria, Lai, Tiziana, Norenburg, Jon L., Curini-Galletti, Marco and Casu, Marco 2015. An 18S and 28S-based clock calibration for marine Proseriata (Platyhelminthes). Journal of experimental marine biology and ecology, 463: 22-31. doi:10.1016/j.jembe.2014.10.020

    Leasi, Francesca and Norenburg, Jon L. 2014. The Necessity of DNA Taxonomy to Reveal Cryptic Diversity and Spatial Distribution of Meiofauna, with a Focus on Nemertea. PLoS ONE, 9(8): 1-15. doi:10.1371/journal.pone.0104385

    Fonseca, Gustavo, Norenburg, Jon L. and Domenico, Maikon Di 2014. Editorial: diversity of marine meiofauna on the coast of Brazil. Marine Biodiversity, : 1-4. doi:10.1007/s12526-014-0261-0

    Andrade, Sonia C. S., Montenegro, Horácio, Strand, Malin, Schwartz, Megan, Kajihara, Hiroshi, Norenburg, Jon L., Turbeville, James M., Sundberg, Per and Giribet, Gonzalo 2014. A transcriptomic approach to ribbon worm systematics (Nemertea): resolving the Pilidiophora problem. Molecular biology and evolution, 31(12): 3206-3215. doi:10.1093/molbev/msu253

    Gonzalez-Cueto, Jaime, Quiroga, Sigmer and Norenburg, Jon L. 2014. A shore-based preliminary survey of marine ribbon worms (Nemertea) from the Caribbean coast of Colombia. Zookeys, 439: 83-108. doi:10.3897/zookeys.439.5965

    Chen, Haixia, Sun, Shichun, Norenburg, Jon L. and Sundberg, Per 2014. Mutation and selection cause codon usage and bias in mitochondrial genomes of ribbon worms. PLoS One, 9(1): 1-11. doi:10.1371/journal.pone.0085631

    Full Publication List

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