Robert HershlerResearch Zoologist

Liu, Hsiu-Ping, Hershler, Robert, Lang, Brian and Davies, Justin 2013. Molecular evidence for cryptic species in a narrowly endemic western North American springsnail (Pyrgulopsis gilae). Conservation Genetics, doi:10.1007/s10592-013-0483-x

Wilke, Thomas, Haase, Martin, Hershler, Robert, Liu, Hsiu-Ping, Misof, Bernhard and Ponder, Winston 2013. Pushing short DNA fragments to the limit: Phylogenetic relationships of 'hydrobioid' gastropods (Caenogastropoda: Rissooidea). Molecular Phylogenetics and Evolution, 66(3): 715-736. doi:10.1016/j.ympev.2012.10.025

Liu, Hsiu-Ping, Walsh, Jessica and Hershler, Robert 2013. taxonomic clarification and phylogeography of Fluminicola coloradensis Morrison, a widely ranging western North American pebblesnail. Monographs of the Western North American Naturalist, 6: 87-110.

Hershler, Robert and Liu, Hsiu-Ping 2012. A new species of springsnail (Pyrgulopsis) from the Owyhee River basin, Nevada. Western North American Naturalist, 72(1): 21-31.

Kabat, Alan R., Hershler, Robert and González-Guillén, Adrian 2012. Resolution of taxonomic problems associated with the complex publication history of the seminal Torre and Bartsch monograph on Cuban Urocoptidae (Gastropoda: Pulmonata). Zootaxa, 3362: 43-53.

Liu, Hsiu-Ping and Hershler, Robert 2012. Phylogeography of an endangered western North American springsnail. Conservation Genetics, 13(1): 299-305. doi:10.1007/s10592-011-0288-8

Liu, H. P., Hershler, Robert, Marn, J. and Worsfold, T. M. 2012. Microsatellite evidence for Tetraploidy in invasive populations of the New Zealand mudsnail, Potamopyrgus antipodarum (Gray, 1843). Journal of Molluscan Studies, 78(2): 227-230. doi:10.1093/mollus/eyr055

Kroll, Oliver, Hershler, Robert, Albrecht, Christian, Terrazas, Edmundo M., Apaza, Roberto, Fuentealba, Carmen, Wolff, Christian and Wilke, Thomas 2012. The endemic gastropod fauna of Lake Titicaca: correlation between molecular evolution and hydrographic history. Ecology and Evolution, 2(7): 1517-1530. doi:10.1002/ece3.280

Hershler, Robert, Liu, Hsiu-Ping and Clark, William H. 2012. Comments on ''The absence of genotypic diversity in a successful parthenogenetic invader'' by Mark Dybdahl and Devin Drown. Biological Invasions, 14(8): 1643-1645. doi:10.1007/s10530-012-0184-2

Hershler, Robert and Liu, Hsiu-Ping 2012. Molecular phylogeny of the western North American pebblesnails, genus Fluminicola (Rissooidea: Lithoglyphidae), with description of a new species. Journal of Molluscan Studies, 78(4): 321-329. doi:10.1093/mollus/eys017

Hershler, Robert, Thompson, Fred G. and Liu, Hsiu-Ping 2011. A large range extension and molecular phylogenetic analysis of the monotypic North American aquatic gastropod genus Cincinnatia (Hydrobiidae). Journal of Molluscan Studies, 77(3): 232-240. doi:10.1093/mollus/eyr008

Hershler, Robert and Liu, Hsiu-Ping 2011. Redescription of Marstonia comalensis (Pilsbry & Ferriss, 1906), a poorly known and possibly threatened freshwater gastropod from the Edwards Plateau region (Texas). ZooKeys, 77: 1-16.

Hershler, Robert and Liu, Hsiu-Ping 2011. Comments on the paper “Is the track of the Yellowstone hotspot driven by a deep mantle plume? — Review of volcanism, faulting, and uplift in light of new data” by Kenneth Pierce and Lisa Morgan [J. Volcanol. Geotherm. Res. 188 (2009), 1–25]. Journal of Volcanology and Geothermal Research, 202(3-4): 262-263.

Hershler, Robert, Liu, Hsiu-Ping and Landye, J. Jerry 2011. New species and records of springsnails (Caenogastropoda: Cochliopidae: Tryonia) from the Chihuahuan Desert (Mexico and United States), an imperiled biodiversity hotspot. Zootaxa, 3001: 1-32.

Hershler, Robert, Liu, Hsiu-Ping and Landye, J. Jerry 2011. Two new genera and four new species of freshwater cochliopid gastropods (Rissooidea) from northeastern Mexico. Journal of Molluscan Studies, 77(1): 8-23.

Hershler, Robert and Liu, Hsiu-Ping 2010. Two new, possibly threatened species of Pyrgulopsis (Gastropoda: Hydrobiidae) from southwestern California. Zootaxa, 2343: 1-17.

Hershler, Robert 2010. Microsatellite evidence of invasion and rapid spread of divergent New Zealand mudsnail (Potamopyrgus antipodarum) clones in the Snake River basin, Idaho, USA. Biological Invasions, 12(6): 1521-1532.

Hershler, Robert, Liu, Hsiu-Ping and Lang, Brian K. 2010. Transfer of Cochliopa texana to Pyrgulopsis (Hydrobiidae) and description of a third congener from the lower Pecos River basin. Journal of Molluscan Studies, 76(3): 245-256.

Hershler, Robert and Jayko, Angela S. 2009. A mactrid bivalve from Pleistocene deposits of Lake Russell, Mono Basin, California. Journal of Paleontology, 83(3): 496-499.

Hershler, Robert and Liu, Hsiu-Ping 2009. New species and records of Pyrgulopsis (Gastropoda: Hydrobiidae) from the Snake River basin, southeastern Oregon: further delineation of a highly imperiled fauna. Zootaxa, 2006: 1-22.

Liu, Hsiu-Ping and Hershler, Robert 2009. Genetic diversity and population structure of the threatened Bliss Rapids snail (Taylorconcha serpenticola). Freshwater Biology, 54(6): 1285-1299.

Hershler, Robert and Liu, Hsiu-Ping 2008. Ancient vicariance and recent dispersal of springsnails (Hydrobiidae: Pyrgulopsis) in the Death Valley system, California-Nevada. In: Reheis, M. C., Hershler, Robert and Miller, D. M., Late Cenozoic drainage history of the southwestern Great Basin and lower Colorado River region: geologic and biologic perspectives. Denver, CO: Geological Society of America Special Paper 439, pp.91-101.

Liu, Hsiu-Ping and Hershler, Robert 2008. Microsatellite markers for the threatened Bliss Rapids snail (Taylorconcha serpenticola) and cross-amplification in its congener, T. inspirata. Molecular Ecology Resources, 8(2): 418-420.

Hershler, Robert and Liu, Hsiu-Ping 2008. Phylogenetic relationships of Assimineid gastropods of the Death Valley-lower Colorado River region: relicts of a late Neogene marine incursion?. Journal of Biogeography, 35(10): 1816-1825.

Hershler, Robert, Liu, Hsiu-Ping and Gustafson, Daniel L. 2008. A second species of Pyrgulopsis (Hydrobiidae) from the Missouri River basin, with molecular evidence supporting faunal origin through Pliocene stream capture across the northern continental divide. Journal of Molluscan Studies, 74(4): 403-413.

Reheis, M. C., Hershler, Robert and Miller, D. M. 2008. Late Cenozoic drainage history of the southwestern Great Basin and lower Colorado River region: geologic and biologic perspectives Denver, CO: Geological Society of America.

Hershler, Robert, Davis, C. L., Kitting, C. L. and Liu, Hsiu-Ping 2007. Discovery of introduced and cryptogenic cochliopid gastropods in the San Francisco Estuary, California. Journal of Molluscan Studies, 73(4): 323-332.

Hershler, Robert, Liu, Hsiu-Ping, Frest, T. J. and Johannes, E. J. 2007. Extensive diversification of pebblesnails (Lithoglyphidae: Fluminicola) in the upper Sacramento River basin. Zoological Journal of the Linnean Society, 149(3): 371-422.

Hershler, Robert, Liu, Hsiu-Ping and Lang, B. K. 2007. Genetic and morphologic variation of the Pecos assiminea, an endangered mollusk of the Rio Grande region, United States and Mexico (Caenogastropoda: Rissooidea: Assimineidae). Hydrobiologia, 579(1): 317-335.

Liu, Hsiu-Ping and Hershler, Robert 2007. A test of the vicariance hypothesis of western North American freshwater biogeography. Journal of Biogeography, 34(3): 534-548.

Hershler, Robert, Liu, P. and Sada, D. W. 2007. Origin and diversification of the Soldier Meadow springsnails (Hydrobiidae: Pyrgulopsis), a species flock in the northwestern Great Basin, United States. Journal of Molluscan Studies, 73(2): 167-183.

Hershler, Robert, Liu, P., Frest, T. J., Johannes, E. J. and Clark, W. H. 2006. Genetic structure of the western North American aquatic gastropod genus Taylorconcha and description of a second species. Journal of Molluscan Studies, 72(2): 167-177.

Hershler, Robert, Mulvey, M. and Liu, P. 2005. Genetic variation in the Desert Springsnail (Tryonia porrecta): implications for reproductive mode and dispersal. Molecular Ecology, 14: 1755-1765.

Liu, Hsiu-Ping and Hershler, Robert 2005. Molecular systematics and radiation of western North American nymphophiline gastropods. Molecular Phylogenetics and Evolution, 34(2): 284-298.

Lydeard, Charles, Cowie, R. H., Bogan, A. E., Bouchet, Philippe, Cummings, K. S., Frest, T. J., Herbert, D. G., Hershler, Robert, Gargominy, O., Perez, K., Ponder, W. F., Roth, B., Seddon, M., Strong, Ellen E. and Thompson, F. G. 2004. The Global Decline of Nonmarine Mollusks. Bioscience, 54: 321-330.

Hershler, Robert and Liu, Hsiu-Ping 2004. A molecular phylogeny of aquatic gastropods provides a new perspective on biogeographic history of the Snake River region. Molecular Phylogenetics and Evolution, 32: 927-937.

Hershler, Robert and Liu, P. 2004. Taxonomic reappraisal of species assisgned to the North American freshwater gastropod subgenus Natricola (Rissooidea: Hydrobiidae). The Veliger, 47(1): 66-81.

Liu, Hsiu-Ping, Hershler, Robert and Clift, K. 2003. Mitochondrial DNA sequences reveal extensive cryptic diversity within a western American springsnail. Molecular Ecology, 12: 2771-2782.

Hershler, Robert, Frest, T. J., Liu, P. and Johannes, E. J. 2003. Rissooidean snails from the Pit River basin, California. The Veliger, 46(4): 275-304.

Hershler, Robert, Liu, P. and Thompson, F. G. 2003. Phylogenetic relationships of North American nymphophiline gastropods based on mitochondrial DNA sequences. Zoologica Scripta, 32: 357-366.

Hershler, Robert, Liu, Hsiu-Ping and Landye, J. J. 2002. A new species of Eremopyprgus (Hydrobiidae: Cochliopinae) from the Chihuahuan desert, Mexico: Phylogentic relationships and biogeography. Journal of Molluscan Studies, 68: 7-13.

Thompson, F. G. and Hershler, Robert 2002. Tepalcatia, a new genus of hydrobiid snails (Prosobranchia: Rissooidea) from the Rio Balsas basin, central Mexico. Proceedings of the Biological Society of Washington, 115(1): 189-204.

Hershler, Robert, Liu, Hsiu-Ping and Stockwell, C. A. 2002. A new genus and species of aquatic gastropods (Rissooidea: Hydrobiidae) from the North American southwest: phylogenetic relationships and biogeography. Proceedings of the Biological Society of Washington, 115: 171-188.

Hershler, Robert, Madsen, D. B. and Currey, D. R. 2002. Great Basin Aquatic Systems History. Smithsonian Contributions to the Earth Sciences, 33: 1-405. doi:10.5479/si.00810274.33.1

Thompson, F. G. and Hershler, Robert 2002. Two genera of North American freshwater snails: Marstonia Baker, 1926, resurrected to generic status, and Flordobia, new Genus (Prosobranchia: Hydrobiidae: Nymphophillinae). The Veliger, 45(3): 269-271.

Hershler, Robert and Gustafson, D. L. 2001. First record for springsnails (Mollusca: Hydrobiidae: Pyrgulopsis) from the northern Rocky Mountains. Proceedings of the Biological Society of Washington, 114(1): 297-308.

Hershler, Robert 2001. Systematics of the North and Central American aquatic snail genus Tryonia (Rissooidea : Hydrobiidae). Smithsonian Contributions to Zoology, 612: 1-53. doi:10.5479/si.00810282.612

Liu, Hsiu-Ping, Hershler, Robert and Thompson, F. G. 2001. Phylogenetic relationships of the Cochliopinae (Rissooidea: Hydrobiidae): an enigmatic group of aquatic gastropods. Molecular Phylogenetics and Evolution, 21(1): 17-25.

Hershler, Robert and Sada, D. W. 2000. A new species of hydrobiid snail of the genus Pyrgulopsis from northwestern Nevada. The Veliger, 43(4): 367-375.

Hershler, Robert, Liu, Hsiu-Ping and Mulvey, M. 1999. Phylogenetic relationships within the aquatic snail genus Tryonia: implications for biogeography of the North American Southwest. Molecular Phylogenetics and Evolution, 13: 377-391.

Hershler, Robert 1999. A systematic review of the hydrobiid snails (Gastropoda: Rissoidea) of the Great Basin, western United States. Part II. Genera Colligyrus, Fluminicola, Pristinicola, and Tryonia. The Veliger, 42(4): 306-337.

Hershler, Robert, Mulvey, M. and Liu, Hsiu-Ping 1999. Biogeography in the Death Valley region: evidence from springsnails (Hydrobiidae: Tryonia). Zoological Journal of the Linnean Society, 126: 335-354.

Hershler, Robert and Ponder, W. F. 1998. A review of morphological characters of hydrobioid snails. Smithsonian Contributions to Zoology, 600: 1-55. doi:10.5479/si.00810282.600

Hershler, Robert 1998. A systematic review of the hydrobiid snails (Gastropoda: Rissoidea) of the Great Basin, western United States. Part I. Genus Pyrgulopsis. The Veliger, 41(1): 1-132.

Hershler, Robert and Thompson, F. G. 1996. Redescription of Paludina integra Say, 1821, type species of Genus Cincinnatia (Gastropoda: Hydrobiidae). Journal of Molluscan Studies, 62: 33-55.

Hershler, Robert and Frest, T. J. 1996. A review of the North American freshwater snail Genus Fluminicola (Hydrobiidae). Smithsonian Contributions to Zoology, 583: 1-41. doi:10.5479/si.00810282.583

Hershler, Robert 1996. Review of the North American aquatic snail genus Probythinella (Rissooidea: Hydrobiidae). Invertebrate Biology, 115(2): 120-144.

Hershler, Robert 1995. New freshwater snails of the genus Pyrgulopsis (Rissooidea: Hydrobiidae) from California. Veliger, 38(4): 343-373.

Hershler, Robert, Frest, T. J., Johannes, E. J., Bowler, P. A. and Thompson, F. G. 1994. Two New Genera of Hydrobiid Snails (Prosobranchia: Rissooidea) from the Northwestern United States. Veliger, 37: 221-243.

Hershler, Robert 1994. A review of the North American freshwater snail genus Pyrgulopsis (Hydrobiidae). Smithsonian Contributions to Zoology, 554: 1-115. doi:10.5479/si.00810282.554

Hershler, Robert and Velkovrh, F. 1993. A New Genus of Hydrobiid Snails (Prosobranchia: Rissooidea) from Northern South America. Proceedings of the Biological Society of Washington, 106: 182-189.

Kabat, Alan R. and Hershler, Robert 1993. The Prosobranch Snail Family Hydrobiidae (Gastropoda: Rissooidea): Review of Classification and Supraspecific Taxa. Smithsonian Contributions to Zoology, 547: 1-94. doi:10.5479/si.00810282.547

Hershler, Robert and Thompson, F. G. 1992. A Review of the Aquatic Gastropod Subfamily Cochliopinae (Prosobranchia: Hydrobiidae). Malacological Review, Supplement, 5: 1-140.

Thompson, F. G. and Hershler, Robert 1991. New Hydrobiid Snails (Mollusca: Gastropoda: Prosobranchia: Truncatelloidea) from North America. Proceedings of the Biological Society of Washington, 104: 669-683.

Thompson, F. G. and Hershler, Robert 1991. Two New Hydrobiid Snails (Amnicolinae) from Florida and Georgia, with a Discussion of the Biogeography of Freshwater Gastropods of South Georgia Streams. Malacological Review, 24: 55-72.

Hershler, Robert and Pratt, W. L. 1990. A New Pyrgulopsis (Gastropoda: Hydrobiidae) from Southeastern California, with a Model for Historical Development of the Death Valley Hydrographic System. Proceedings of the Biological Society of Washington, 103: 279-299.

Hershler, Robert, Pierson, J. M. and Krotzer, R. S. 1990. Rediscovery of Tulotoma magnifica (Conrad)(Gastropoda: Viviparidae). Proceedings of the Biological Society of Washington, 103: 815-824.

Hershler, Robert 1990. Pyrgulopsis bruneauensis, a New Springsnail (Gastropoda: Hydrobiidae) from the Snake River Plain, Southern Idaho. Proceedings of the Biological Society of Washington, 103: 803-814.

Hershler, Robert and Thompson, F. G. 1990. Antrorbis breweri, a new Genus and Species of hydrobiid cave snail from Coosa River Basin, Northeastern Alabama. Proceedings of the Biological Society of Washington, 103: 197-204.

Hershler, Robert, Holsinger, John R. and Hubricht, L. 1990. A Revision of the North American Freshwater Snail Genus Fontigens (Prosobranchia: Hydrobiidae). Smithsonian Contributions to Zoology, 509: 1-49. doi:10.5479/si.00810282.509

Hershler, Robert and Holsinger, John R. 1990. Zoogeography of North American Hydrobiid Cavesnails. Stygologia, 5: 5-16.

Ponder, W. F., Hershler, Robert and Jenkins, B. J. 1989. An Endemic Radiation of Hydrobiid Snails from Artesian Springs in Northern South Australia: Their Taxonomy, Physiology, Distribution and Anatomy. Malacologia, 31: 1-140.

Hershler, Robert 1989. Springsnails (Gastropoda: Hydrobiidae) of Owens and Amargosa River (Exclusive of Ash Meadows) Drainages, Death Valley System, California-Nevada. Proceedings of the Biological Society of Washington, 102: 176-248.

Hershler, Robert 1989. Holsingeria unthanksensis, a new genus and species of aquatic cavesnail from eastern North America. Malacological Review, 22: 93-100.

Hershler, Robert and Thompson, F. G. 1988. Notes on Morphology of Amnicola limosa (Say, 1817)(Gastropoda: Hydrobiidae) with Comments on Status of the Subfamily Amnicolinae. Malacological Review, 22: 81-92.

Hershler, Robert 1988. Holsingeria unthanksensis, a New Genus and Species of Aquatic Cavesnail from Eastern North America. Malacological Review, 22: 93-100.

Hershler, Robert and Hayek, Lee-Ann C. 1988. Shell variation of springsnail populations in the Cuatro Ciénegas basin, Mexico: preliminary analysis of limnocrene fauna. The Nautilus, 102: 56-64.

Hershler, Robert and Hubricht, L. 1988. Notes on Antroselates Hubricht, 1963 and Antrobia Hubricht, 1971 (Gastropoda: Hydrobiidae). Proceedings of the Biological Society of Washington, 101: 730-740.

Hershler, Robert and Landye, J. 1988. Arizona Hydrobiidae. Smithsonian Contributions to Zoology, 459: 1-63. doi:10.5479/si.00810282.459

Hershler, Robert 1987. Redescription of Assiminea infima Berry, 1947 from Death Valley, California. Veliger, 29: 274-288.

Hershler, Robert and Sada, D. 1987. Springsnails (Gastropoda: Hydrobiidae) of Ash Meadows, Amargosa Basin, California-Nevada. Proceedings of the Biological Society of Washington, 100: 776-843.

Hershler, Robert and Longley, G. 1987. Phreatoceras, a New Name for Hadoceras Hershler and Longley, 1986 (Gastropoda) non Strand, 1934 (Cephalopoda). Proceedings of the Biological Society of Washington, 100: 402

Hershler, Robert and Longley, G. 1987. Phreatodrobia coronae, a New Species of Cavesnail from Southwestern Texas. The Nautilus, 101: 133-139.

Hershler, Robert and Thompson, F. G. 1987. North American Hydrobiidae (Gastropoda: Rissoacea): Redescription and Systematic Relationships of Tryonia Stimpson, 1865 and Pyrgulopsis Call and Pilsbry, 1886. The Nautilus, 101: 25-32.

Hershler, Robert and Minckley, W. L. 1986. Microgeographic Variation in the Banded Spring Snail (Hydrobiidae: Mexipyrgus) from the Cuatro Cienegas Basin, Coahuila, Mexico. Malacologia, 27: 357-374.

Hershler, Robert and Longley, G. 1986. Phreatic Hydrobiids (Gastropoda: Prosobranchia) from the Edwards (Balcones Fault Zone) Aquifer Region, South-Central Texas. Malacologia, 27: 127-172.

Hershler, Robert and Longley, G. 1986. Hadoceras taylori, a New Genus and Species of Phreatic Hydrobiidae (Gastropoda: Rissoacea) from South-central Texas. Proceedings of the Biological Society of Washington, 99: 121-136.

Hershler, Robert 1985. Systematic Revision of the Hydrobiidae (Gastropoda: Rissoacea) of the Cuatro Cienegas Basin, Coahuila, Mexico. Malacologia, 26: 31-123.

Hershler, Robert 1984. Hydrobiid Snails of the Cuatro Cienegas Basin, Mexico: Systematic Relationships and Ecology of a Unique Fauna. Journal of the Arizona-Nevada Academy of Science, 19: 61-76.

Hershler, Robert and Davis, G. M. 1980. The Morphology of Hydrobia truncata: Relevance to Systematics of Hydrobia. Biological Bulletin, 158: 195-219.